1.Key Laboratory of Exploration and Utilization of Aquatic Resources, Ministry of Education, Shanghai Ocean University, Shanghai 201306, China

2.Shanghai Engineering Research Center of Aquaculture, Shanghai Ocean University, Shanghai 201306, China

3.Department of Marine Organism Taxonomy and Phylogeny, Institute of Oceanology, Chinese Academy of Sciences(IOCAS), Qingdao 266071, China

4.Key Laboratory of Marine Ecology and Environmental Sciences, Institute of Oceanology, Chinese Academy of Sciences, Qingdao 266071, China